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Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on

most body segments; they are known scientifically as the class Diplopoda, the name being derived
from this feature. Each double-legged segment is a result of two single segments fused together.
Most millipedes have very elongated cylindrical or flattened bodies with more than 20 segments,
while pill millipedes are shorter and can roll into a ball. Although the name "millipede" derives from
the Latin for "thousand feet", no known species has 1,000; the record of 750 legs belongs to Illacme
plenipes. There are approximately 12,000 named species classified into 16 orders and around
140 families, making Diplopoda the largest class of myriapods, an arthropod group which also
includes centipedes and other multi-legged creatures.
Most millipedes are slow-moving detritivores, eating decaying leaves and other dead plant matter.
Some eat fungi or suck plant fluids, and a small minority are predatory. Millipedes are generally
harmless to humans, although some can become household or garden pests, especially
in greenhouses where they can cause severe damage to emergent seedlings. Most millipedes
defend themselves with a variety of chemicals secreted from pores along the body, although the
tiny bristle millipedes are covered with tufts of detachable bristles. Reproduction in most species is
carried out by modified male legs called gonopods, which transfer packets of sperm to females.
First appearing in the Silurian period, millipedes are some of the oldest known land animals. Some
members of prehistoric groups grew to over 2 m (6 ft 7 in); the largest modern species reach
maximum lengths of 27 to 38 cm (11 to 15 in). The longest extant species is the giant African
millipede (Archispirostreptus gigas).
Among myriapods, millipedes have traditionally been considered most closely related to the
tiny pauropods, although some molecular studies challenge this relationship. Millipedes can be
distinguished from the somewhat similar but only distantly related centipedes (class Chilopoda),
which move rapidly, are carnivorous, and have only a single pair of legs on each body segment. The
scientific study of millipedes is known as diplopodology, and a scientist who studies them is called a
diplopodologist.

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